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In most muscles, all the fibers are oriented in the same direction, running in a line from the origin to the insertion. However, In pennate muscles, the individual fibers are oriented at an angle relative to the line of action, attaching to the origin and insertion tendons at each end.

Because the contracting fibers are pulling at an angle to the overall action of the muscle, the change in length is smaller, but this same orientation allows for more fibers thus more force in a muscle of a given size.

Pennate muscles are usually found where their length change is less important than maximum force, such as the rectus femoris. Skeletal muscle is arranged in discrete muscles, an example of which is the biceps brachii biceps.

The tough, fibrous epimysium of skeletal muscle is both connected to and continuous with the tendons. In turn, the tendons connect to the periosteum layer surrounding the bones, permitting the transfer of force from the muscles to the skeleton.

Together, these fibrous layers, along with tendons and ligaments, constitute the deep fascia of the body. The muscular system consists of all the muscles present in a single body.

There are approximately skeletal muscles in the human body, [14] but an exact number is difficult to define. The difficulty lies partly in the fact that different sources group the muscles differently and partly in that some muscles, such as palmaris longus , are not always present.

A muscular slip is a narrow length of muscle that acts to augment a larger muscle or muscles. The muscular system is one component of the musculoskeletal system , which includes not only the muscles but also the bones, joints, tendons, and other structures that permit movement.

All muscles are derived from paraxial mesoderm. The paraxial mesoderm is divided along the embryo 's length into somites , corresponding to the segmentation of the body most obviously seen in the vertebral column.

The myotome is divided into two sections, the epimere and hypomere, which form epaxial and hypaxial muscles , respectively. The only epaxial muscles in humans are the erector spinae and small intervertebral muscles, and are innervated by the dorsal rami of the spinal nerves.

All other muscles, including those of the limbs are hypaxial, and inervated by the ventral rami of the spinal nerves. During development, myoblasts muscle progenitor cells either remain in the somite to form muscles associated with the vertebral column or migrate out into the body to form all other muscles.

Myoblast migration is preceded by the formation of connective tissue frameworks, usually formed from the somatic lateral plate mesoderm.

Myoblasts follow chemical signals to the appropriate locations, where they fuse into elongate skeletal muscle cells. The three types of muscle skeletal, cardiac and smooth have significant differences.

However, all three use the movement of actin against myosin to create contraction. In skeletal muscle, contraction is stimulated by electrical impulses transmitted by the nerves , the motoneurons motor nerves in particular.

Cardiac and smooth muscle contractions are stimulated by internal pacemaker cells which regularly contract, and propagate contractions to other muscle cells they are in contact with.

All skeletal muscle and many smooth muscle contractions are facilitated by the neurotransmitter acetylcholine.

The action a muscle generates is determined by the origin and insertion locations. The cross-sectional area of a muscle rather than volume or length determines the amount of force it can generate by defining the number of "sarcomeres" which can operate in parallel.

Each skeletal muscle contains long units called myofibrils, and each myofibril is a chain of sarcomeres.

Since contraction occurs at the same time for all connected sarcomeres in a muscles cell, these chains of sarcomeres shorten together, thus shortening the muscle fiber, resulting in overall length change.

For example, moving the insertion point of the biceps more distally on the radius farther from the joint of rotation would increase the force generated during flexion and, as a result, the maximum weight lifted in this movement , but decrease the maximum speed of flexion.

Moving the insertion point proximally closer to the joint of rotation would result in decreased force but increased velocity.

This can be most easily seen by comparing the limb of a mole to a horse—in the former, the insertion point is positioned to maximize force for digging , while in the latter, the insertion point is positioned to maximize speed for running.

The efferent leg of the peripheral nervous system is responsible for conveying commands to the muscles and glands, and is ultimately responsible for voluntary movement.

Nerves move muscles in response to voluntary and autonomic involuntary signals from the brain. Deep muscles, superficial muscles, muscles of the face and internal muscles all correspond with dedicated regions in the primary motor cortex of the brain , directly anterior to the central sulcus that divides the frontal and parietal lobes.

In addition, muscles react to reflexive nerve stimuli that do not always send signals all the way to the brain. In this case, the signal from the afferent fiber does not reach the brain, but produces the reflexive movement by direct connections with the efferent nerves in the spine.

However, the majority of muscle activity is volitional, and the result of complex interactions between various areas of the brain. Nerves that control skeletal muscles in mammals correspond with neuron groups along the primary motor cortex of the brain's cerebral cortex.

Commands are routed through the basal ganglia and are modified by input from the cerebellum before being relayed through the pyramidal tract to the spinal cord and from there to the motor end plate at the muscles.

Along the way, feedback, such as that of the extrapyramidal system contribute signals to influence muscle tone and response.

Deeper muscles such as those involved in posture often are controlled from nuclei in the brain stem and basal ganglia.

In skeletal muscles, muscle spindles convey information about the degree of muscle length and stretch to the central nervous system to assist in maintaining posture and joint position.

The sense of where our bodies are in space is called proprioception , the perception of body awareness, the "unconscious" awareness of where the various regions of the body are located at any one time.

Several areas in the brain coordinate movement and position with the feedback information gained from proprioception.

The cerebellum and red nucleus in particular continuously sample position against movement and make minor corrections to assure smooth motion.

Muscular activity accounts for much of the body's energy consumption. All muscle cells produce adenosine triphosphate ATP molecules which are used to power the movement of the myosin heads.

Muscles have a short-term store of energy in the form of creatine phosphate which is generated from ATP and can regenerate ATP when needed with creatine kinase.

Muscles also keep a storage form of glucose in the form of glycogen. Glycogen can be rapidly converted to glucose when energy is required for sustained, powerful contractions.

Within the voluntary skeletal muscles, the glucose molecule can be metabolized anaerobically in a process called glycolysis which produces two ATP and two lactic acid molecules in the process note that in aerobic conditions, lactate is not formed; instead pyruvate is formed and transmitted through the citric acid cycle.

Muscle cells also contain globules of fat, which are used for energy during aerobic exercise. The aerobic energy systems take longer to produce the ATP and reach peak efficiency, and requires many more biochemical steps, but produces significantly more ATP than anaerobic glycolysis.

Cardiac muscle on the other hand, can readily consume any of the three macronutrients protein, glucose and fat aerobically without a 'warm up' period and always extracts the maximum ATP yield from any molecule involved.

The heart, liver and red blood cells will also consume lactic acid produced and excreted by skeletal muscles during exercise.

At rest, skeletal muscle consumes This is larger than adipose tissue fat at The efficiency is defined as the ratio of mechanical work output to the total metabolic cost, as can be calculated from oxygen consumption.

The latter two losses are dependent on the type of exercise and the type of muscle fibers being used fast-twitch or slow-twitch.

For an overall efficiency of 20 percent, one watt of mechanical power is equivalent to 4. For example, one manufacturer of rowing equipment calibrates its rowing ergometer to count burned calories as equal to four times the actual mechanical work, plus kcal per hour, [18] this amounts to about 20 percent efficiency at watts of mechanical output.

These can be synthesized experimentally using work loop analysis. Muscle is a result of three factors that overlap: physiological strength muscle size, cross sectional area, available crossbridging, responses to training , neurological strength how strong or weak is the signal that tells the muscle to contract , and mechanical strength muscle's force angle on the lever, moment arm length, joint capabilities.

The force generated by a contraction can be measured non-invasively using either mechanomyography or phonomyography , be measured in vivo using tendon strain if a prominent tendon is present , or be measured directly using more invasive methods.

The strength of any given muscle, in terms of force exerted on the skeleton, depends upon length, shortening speed , cross sectional area, pennation , sarcomere length, myosin isoforms, and neural activation of motor units.

Significant reductions in muscle strength can indicate underlying pathology, with the chart at right used as a guide. Since three factors affect muscular strength simultaneously and muscles never work individually, it is misleading to compare strength in individual muscles, and state that one is the "strongest".

But below are several muscles whose strength is noteworthy for different reasons. Exercise is often recommended as a means of improving motor skills , fitness , muscle and bone strength, and joint function.

Exercise has several effects upon muscles, connective tissue , bone, and the nerves that stimulate the muscles. One such effect is muscle hypertrophy , an increase in size of muscle due to an increase in the number of muscle fibers or cross-sectional area of myofibrils.

Generally, there are two types of exercise regimes, aerobic and anaerobic. Aerobic exercise e. Aerobic activities rely on the aerobic respiration i.

Muscles involved in aerobic exercises contain a higher percentage of Type I or slow-twitch muscle fibers, which primarily contain mitochondrial and oxidation enzymes associated with aerobic respiration.

The anaerobic activities predominately use Type II, fast-twitch, muscle fibers. Many exercises are partially aerobic and anaerobic; for example, soccer and rock climbing.

The presence of lactic acid has an inhibitory effect on ATP generation within the muscle. It can even stop ATP production if the intracellular concentration becomes too high.

However, endurance training mitigates the buildup of lactic acid through increased capillarization and myoglobin. Once moved out of muscles, lactic acid can be used by other muscles or body tissues as a source of energy, or transported to the liver where it is converted back to pyruvate.

In addition to increasing the level of lactic acid, strenuous exercise results in the loss of potassium ions in muscle. This may facilitate the recovery of muscle function by protecting against fatigue.

Delayed onset muscle soreness is pain or discomfort that may be felt one to three days after exercising and generally subsides two to three days after which.

Once thought to be caused by lactic acid build-up, a more recent theory is that it is caused by tiny tears in the muscle fibers caused by eccentric contraction , or unaccustomed training levels.

Since lactic acid disperses fairly rapidly, it could not explain pain experienced days after exercise. Independent of strength and performance measures, muscles can be induced to grow larger by a number of factors, including hormone signaling, developmental factors, strength training , and disease.

Contrary to popular belief, the number of muscle fibres cannot be increased through exercise. Instead, muscles grow larger through a combination of muscle cell growth as new protein filaments are added along with additional mass provided by undifferentiated satellite cells alongside the existing muscle cells.

Biological factors such as age and hormone levels can affect muscle hypertrophy. During puberty in males, hypertrophy occurs at an accelerated rate as the levels of growth-stimulating hormones produced by the body increase.

Natural hypertrophy normally stops at full growth in the late teens. As testosterone is one of the body's major growth hormones, on average, men find hypertrophy much easier to achieve than women.

Taking additional testosterone or other anabolic steroids will increase muscular hypertrophy. Muscular, spinal and neural factors all affect muscle building.

Sometimes a person may notice an increase in strength in a given muscle even though only its opposite has been subject to exercise, such as when a bodybuilder finds her left biceps stronger after completing a regimen focusing only on the right biceps.

This phenomenon is called cross education. During ordinary living activities, between 1 and 2 percent of muscle is broken down and rebuilt daily. Inactivity and starvation in mammals lead to atrophy of skeletal muscle, a decrease in muscle mass that may be accompanied by a smaller number and size of the muscle cells as well as lower protein content.

In humans, prolonged periods of immobilization, as in the cases of bed rest or astronauts flying in space, are known to result in muscle weakening and atrophy.

During aging, there is a gradual decrease in the ability to maintain skeletal muscle function and mass, known as sarcopenia. The exact cause of sarcopenia is unknown, but it may be due to a combination of the gradual failure in the "satellite cells" that help to regenerate skeletal muscle fibers, and a decrease in sensitivity to or the availability of critical secreted growth factors that are necessary to maintain muscle mass and satellite cell survival.

Sarcopenia is a normal aspect of aging, and is not actually a disease state yet can be linked to many injuries in the elderly population as well as decreasing quality of life.

There are also many diseases and conditions that cause muscle atrophy. Examples include cancer and AIDS , which induce a body wasting syndrome called cachexia.

Other syndromes or conditions that can induce skeletal muscle atrophy are congestive heart disease and some diseases of the liver.

In general, problems with nervous control can cause spasticity or paralysis , depending on the location and nature of the problem.

A large proportion of neurological disorders , ranging from cerebrovascular accident stroke and Parkinson's disease to Creutzfeldt—Jakob disease , can lead to problems with movement or motor coordination.

Symptoms of muscle diseases may include weakness , spasticity, myoclonus and myalgia. Diagnostic procedures that may reveal muscular disorders include testing creatine kinase levels in the blood and electromyography measuring electrical activity in muscles.

In some cases, muscle biopsy may be done to identify a myopathy , as well as genetic testing to identify DNA abnormalities associated with specific myopathies and dystrophies.

A non-invasive elastography technique that measures muscle noise is undergoing experimentation to provide a way of monitoring neuromuscular disease.

The sound produced by a muscle comes from the shortening of actomyosin filaments along the axis of the muscle. During contraction , the muscle shortens along its longitudinal axis and expands across the transverse axis , producing vibrations at the surface.

The evolutionary origin of muscle cells in metazoans is a highly debated topic. In one line of thought scientists have believed that muscle cells evolved once and thus all animals with muscles cells have a single common ancestor.

In the other line of thought, scientists believe muscles cells evolved more than once and any morphological or structural similarities are due to convergent evolution and genes that predate the evolution of muscle and even the mesoderm —the germ layer from which many scientists believe true muscle cells derive.

Schmid and Seipel argue that the origin of muscle cells is a monophyletic trait that occurred concurrently with the development of the digestive and nervous systems of all animals and that this origin can be traced to a single metazoan ancestor in which muscle cells are present.

They argue that molecular and morphological similarities between the muscles cells in cnidaria and ctenophora are similar enough to those of bilaterians that there would be one ancestor in metazoans from which muscle cells derive.

In this case, Schmid and Seipel argue that the last common ancestor of bilateria, ctenophora, and cnidaria was a triploblast or an organism with three germ layers and that diploblasty , meaning an organism with two germ layers, evolved secondarily due to their observation of the lack of mesoderm or muscle found in most cnidarians and ctenophores.

By comparing the morphology of cnidarians and ctenophores to bilaterians, Schmid and Seipel were able to conclude that there were myoblast -like structures in the tentacles and gut of some species of cnidarians and in the tentacles of ctenophores.

Since this is a structure unique to muscle cells, these scientists determined based on the data collected by their peers that this is a marker for striated muscles similar to that observed in bilaterians.

The authors also remark that the muscle cells found in cnidarians and ctenophores are often contests due to the origin of these muscle cells being the ectoderm rather than the mesoderm or mesendoderm.

The origin of true muscles cells is argued by others to be the endoderm portion of the mesoderm and the endoderm.

However, Schmid and Seipel counter this skepticism about whether or not the muscle cells found in ctenophores and cnidarians are true muscle cells by considering that cnidarians develop through a medusa stage and polyp stage.

They observe that in the hydrozoan medusa stage there is a layer of cells that separate from the distal side of the ectoderm to form the striated muscle cells in a way that seems similar to that of the mesoderm and call this third separated layer of cells the ectocodon.

They also argue that not all muscle cells are derived from the mesendoderm in bilaterians with key examples being that in both the eye muscles of vertebrates and the muscles of spiralians these cells derive from the ectodermal mesoderm rather than the endodermal mesoderm.

Furthermore, Schmid and Seipel argue that since myogenesis does occur in cnidarians with the help of molecular regulatory elements found in the specification of muscles cells in bilaterians that there is evidence for a single origin for striated muscle.

In contrast to this argument for a single origin of muscle cells, Steinmetz et al. This author uses an example of the contractile elements present in the porifera or sponges that do truly lack this striated muscle containing this protein.

Furthermore, Steinmetz et al. Steimetz et al. Thus, the usage of any of these structural or regulatory elements in determining whether or not the muscle cells of the cnidarians and ctenophores are similar enough to the muscle cells of the bilaterians to confirm a single lineage is questionable according to Steinmetz et al.

Furthermore, Steinmetz et all showed that the localization of this duplicated set of genes that serve both the function of facilitating the formation of striated muscle genes and cell regulation and movement genes were already separated into striated myhc and non-muscle myhc.

This separation of the duplicated set of genes is shown through the localization of the striated myhc to the contractile vacuole in sponges while the non-muscle myhc was more diffusely expressed during developmental cell shape and change.

Steinmetz et al. Thus, Steinmetz et al. Furthermore, the Z-disc seemed to have evolved differently even within bilaterians and there is a great deal diversity of proteins developed even between this clade, showing a large degree of radiation for muscle cells.

Through this divergence of the Z-disc , Steimetz et al. Through further molecular marker testing, Steinmetz et al.

Through this analysis the authors conclude that due to the lack of elements that bilaterians muscles are dependent on for structure and usage, nonbilaterian muscles must be of a different origin with a different set regulatory and structural proteins.

In another take on the argument, Andrikou and Arnone use the newly available data on gene regulatory networks to look at how the hierarchy of genes and morphogens and other mechanism of tissue specification diverge and are similar among early deuterostomes and protostomes.

By understanding not only what genes are present in all bilaterians but also the time and place of deployment of these genes, Andrikou and Arnone discuss a deeper understanding of the evolution of myogenesis.

In their paper Andrikou and Arnone argue that to truly understand the evolution of muscle cells the function of transcriptional regulators must be understood in the context of other external and internal interactions.

Through their analysis, Andrikou and Arnone found that there were conserved orthologues of the gene regulatory network in both invertebrate bilaterians and in cnidarians.

They argue that having this common, general regulatory circuit allowed for a high degree of divergence from a single well functioning network.

Andrikou and Arnone found that the orthologues of genes found in vertebrates had been changed through different types of structural mutations in the invertebrate deuterostomes and protostomes, and they argue that these structural changes in the genes allowed for a large divergence of muscle function and muscle formation in these species.

Andrikou and Arnone were able to recognize not only any difference due to mutation in the genes found in vertebrates and invertebrates but also the integration of species specific genes that could also cause divergence from the original gene regulatory network function.

Thus, although a common muscle patterning system has been determined, they argue that this could be due to a more ancestral gene regulatory network being coopted several times across lineages with additional genes and mutations causing very divergent development of muscles.

Thus it seems that myogenic patterning framework may be an ancestral trait. However, Andrikou and Arnone explain that the basic muscle patterning structure must also be considered in combination with the cis regulatory elements present at different times during development.

In contrast with the high level of gene family apparatuses structure, Andrikou and Arnone found that the cis regulatory elements were not well conserved both in time and place in the network which could show a large degree of divergence in the formation of muscle cells.

Through this analysis, it seems that the myogenic GRN is an ancestral GRN with actual changes in myogenic function and structure possibly being linked to later coopts of genes at different times and places.

Vertebrate smooth muscle was found to have evolved independently from the skeletal and cardiac muscle types.

From Wikipedia, the free encyclopedia. Contractile soft tissue of mammals. For other uses, see Muscle disambiguation. Main article: Muscle tissue. Main articles: Myocyte and Sarcomere.

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